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An important question that is being increasingly studied across subdisciplines of Heliophysics is “how do mesoscale phenomena contribute to the global response of the system?” This review paper focuses on this question within two specific but interlinked regions in Near-Earth space: the magnetotail’s transition region to the inner magnetosphere and the ionosphere. There is a concerted effort within the Geospace Environment Modeling (GEM) community to understand the degree to which mesoscale transport in the magnetotail contributes to the global dynamics of magnetic flux transport and dipolarization, particle transport and injections contributing to the storm-time ring current development, and the substorm current wedge. Because the magnetosphere-ionosphere is a tightly coupled system, it is also important to understand how mesoscale transport in the magnetotail impacts auroral precipitation and the global ionospheric system response. Groups within the Coupling, Energetics and Dynamics of Atmospheric Regions Program (CEDAR) community have also been studying how the ionosphere-thermosphere responds to these mesoscale drivers. These specific open questions are part of a larger need to better characterize and quantify mesoscale “messengers” or “conduits” of information—magnetic flux, particle flux, current, and energy—which are key to understanding the global system. After reviewing recent progress and open questions, we suggest datasets that, if developed in the future, will help answer these questions. 

Abstract Large changes of the magnetic field associated with magnetic perturbation events (MPEs) with amplitudes |ΔB| of hundreds of nT and 5–10 min duration have been frequently observed within a few hours of midnight. This study compares the statistical location of nighttime MPEs with |dB/dt| ≥ 6 nT/s within the auroral current system observed during 2015 and 2017 at two stations, Cape Dorset and Kuujjuarapik, in Eastern Canada. Maps of the two dimensional nightside auroral current system were derived using the Spherical Elementary Current Systems (SECS) technique. Analyses were produced at each station for all events, and for premidnight and postmidnight subsets. We examine four MPE intervals in detail, two accompanied by auroral images, and show the varying associations between MPEs and overhead ionospheric current systems including electrojets and the field‐aligned like currents. We find 225 of 279 MPEs occurred within the westward electrojet and only 3 within the eastward electrojet. For the premidnight MPEs 100 of 230 events occurred within the Harang current system while many of the remainder occurred within either the downward region 1 current system or the upward region 2 current system. Many of the 49 postmidnight MPEs occurred in either the downward region 1 (11 events) or upward region 2 current system (27 events). These result suggest that the source of MPEs in the premidnight sector is somewhere between the inner to mid plasma sheet and the source for the MPEs in the postmidnight sector is somewhere between the inner magnetosphere and the inner plasma sheet.